Freshwater Fishes of Iran


Acknowledgements

Revised:  26 June 2007

Acknowledgements     Purpose     Materials and Methods     History of Research     Fisheries     Geography     Climate     Habitats     Environmental Change     Drainage Basins     Scientific Names     Fish Structure     Collecting Fishes     Preserving Fishes     Quotes

A wide range of people in Iran, Canada and elsewhere have assisted me in this work over more than 30 years. Inevitably, I will have forgotten some names which I regret. Some people I never met formally, an example being the gentleman nattily dressed in suit by a stream near Kazerun who jumped fully-clothed into the water to help me catch fish. Numerous other Iranians have assisted my studies and this website is dedicated to them.

The staff at the Department of Biology, Shiraz (then Pahlavi) University helped me in numerous ways to collect fishes during a three-year tenure as an Associate Professor. Dr. Bahman Kholdebarin was Chairman of the Department for much of my time in Iran and it is only through his support that I was able to make the collections that enabled this work to be done. The Research Council of Pahlavi University funded field trips and is gratefully acknowledged for this support. Collections were made with the help of drivers and assistants and their efforts over long periods in the field are gratefully acknowledged. They include H. Assadi, M. H. Jaferi, Sh. Mansoorabadi, A. Shirazi, A. Tofangdar and N. Yaghar. Various other people assisted too and are mentioned below under the Pahlavi University name.

Studies on Iranian fishes since my residence in Iran have been supported by grants from the Canadian Museum of Nature, Ottawa (CMN, fish collection acronym CMNFI), by assistance from staff there including Noel Alfonso, Jadwiga Frank, C. G. Gruchy, Sylvie Laframboise, Alison Murray, Claude Renaud and Michèle Steigerwald, and by a wide range of students and volunteers. The staff in the CMN library searched out all the numerous and varied papers on fishes in Iran and neighbouring countries without which this synthesis would not be possible. One paper took six years to locate and arrived in the form of a microfilm from the Soviet Union. I am particularly indebted to Victor Adomaitis who kindly volunteered for the unrewarding task of scanning hundreds of images and converting them to thumbnails and usable files. Mollie MacCormac carried on this task, making a wide variety of images available for the website.

In particular, I should like to acknowledge the support and encouragement of the late Dr. D. E. McAllister, Curator of Fishes, CMN over many years, in terms of training and education, both formal and informal, of financial and moral support, and in practical terms in the ways and means of collecting, cataloguing, identifying, and studying fishes, and of getting things done.

Co-authors are evident in the Bibliography and their added expertise made several studies possible (authors marked by a * are first authors on some papers with me). These are in temporal order F. Krupp, H. F. Alkahem, R. J. Behnke, G. B. Delmastro, M. Kuru, M. Sarieyyüpoglu, F. Papahn, L. A. J..Al-Hassan, A. Abdoli, N. Najafpour, *D. González-Solís, *F. Moravec, Y. Keivany, J. Holcík, J. W. Atz, M. Naderi, *H. Ahnelt, *N. A. Hussein, *T. S. Ali, K. E. Limburg, B. Ya. Vilenkin, N. Keyzer-de Ville, *H. R. Esmaeili, T. T. Nalbant, and *T. Hrbek.

Various people and their organisations are mentioned below separately for their particular assistance; these are in alphabetical order.

Dr. Asghar Abdoli collected numerous specimens including exotics and allowed me to incorporate these discoveries in several papers.

Dr. P. Bănărescu, Institutul de Biologie, Bucureşti has communicated much information in detailed letters on fishes in the Middle East as well as loaning and exchanging specimens, for all of which his assistance is acknowledged.

Dr. R. J. Behnke, Colorado State University, Fort Collins is gratefully acknowledged for his extensive loans of, and access to, collections he and associates made. These are listed more fully in the Materials and Methods.

Prof. Dr. P. G. Bianco, University of Naples, allowed me free access to materials, including types, in his possession at the University of Naples and his hospitality is acknowledged.

Dr. N. Bogutskaya and Dr. A. Naseka, Laboratory of Ichthyology, Zoological Institute, Academy of Sciences, St. Petersburg are thanked especially for their hospitality, access to collections, data analyses and interpretations on Iranian fishes.

Dr. C. E. Bond, Department of Fisheries and Wildlife, Oregon State University, Corvallis allowed extensive loans of fishes from Iran under his care and these materials are listed in the Material and Methods (see Contents).

Staff at the Fish Section, British Museum (Natural History) (now the Natural History Museum) have loaned materials and hosted visits on numerous occasions; their help has been much appreciated for the extensive collections are a required study to understand the Iranian fauna. They include Dr. K. Banister, B. Brewster, P. Campbell, O. Crimmen, S. Davidson, Dr. P. H. Greenwood, A.-M. Hodges, G. Howes, J. Maclaine, Dr. N. Merrett, Dr. D. Siebert, Dr. E. Trewavas, A. Wheeler and Dr. P. J. P. Whitehead.

Dr. T. Hrbek, Washington University School of Medicine, St. Louis is acknowledged for his complementary studies on tooth-carps using molecular techniques.

Dr. M. Kasparek and Prof. Dr. R. Kinzelbach kindly appointed me to the Advisory Board of the journal Zoology in the Middle East which has given me an interesting and valuable overview of studies in that region.

Dr. Yazdan Keivany translated abstracts of his manuscript reports and first posted my bibliography of Iranian freshwater fishes on the internet - a stimulus to this work! He is continuing collaborative efforts aimed at improving knowledge on the ichthyofauna of Iran.

Dr. Bahram Kiabi, Gorgan University of Agricultural Sciences and Natural Resources is thanked for various items of information on fishes, translations and gifts of Farsi articles and many interesting fish specimens. His efforts at facilitating collegiality and his students have formed the core of modern university researchers on the fishes of Iran.

Dr. F. Krupp, Johannes Gutenburg-Universität Mainz and Forschungsintitut Senckenberg (NaturMuseum Senckenberg), Frankfurt am Main contributed a wide variety of information on Middle Eastern fishes, sent me copies of his theses and in his letters provided many stimulating points of discussion which helped me clarify my views on the fishes. His published works are a model for students on fishes in that region. He, with Prof. Dr. Kinzelbach, kindly invited me to the Symposium on the Fauna and Zoogeography of the Middle East in Mainz, 1985.

Nasser Najafpour, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi,Ahvaz was instrumental in arranging visits to Iran and associated field trips. His enthusiastic cooperation in the field resulted in many interesting new specimens and his studies on distributions of fishes in Khuzestan have been very important for this web site. The team at Ahvaz is acknowledged below individually and in teaching me Farsi names of fishes. J. Gh. Marammazi was head of that team and his hospitality and efforts to bring me to Iran are gratefully acknowledged.

Dr. T. T. Nalbant, National Museum of Natural History "Grigore Antipa", Bucharest, is currently studying loaches I collected in Iran.

Staff at the National Museum of Natural History (Smithsonian Institution), Washington arranged loans of specimens and allowed access to the collections during several visits. They include K. A. Bruwelheide, Dr. B. B. Collette, S. Jewett, S. Karnella and Dr. J. T. Williams.

Staff at the Fischsammlung, Naturhistorisches Museum Vienna have also loaned materials and hosted visits and their assistance has been essential to studies on Iranian fishes based on the collections of J. J. Heckel. They include Dr. H. Ahnelt, Dr. E. Mikschi, Dr. B. Herzig and Dr. R. Hacker.

Dr. J. G. Nielsen and Dr. P. R. Möller, Zoological Museum, University of Copenhagen facilitated access to collections despite the "orkan".

Dr. P. Bartsch and Mrs. C. Lamour, Museum für Naturkunde, Humboldt-Universität zu Berlin for access to collections.

M. Rabaniha and F. Owfi, Persian Gulf Fisheries Research Centre, Bushehr and Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Tehran, kindly copied the MMTT catalogue for me and showed me various specimens of fishes from their work in Bushehr Province and southern Iran.

Dr. Jalal Valiallahi provided stimulating discussions on the limits and the content of the genus Barbus in Iran while working at the CMN as well as a variety of photographs of these sometimes immense fish.

Prof. Dr. H. Wilkens, Zoologisches Institut und Zoologisches Museum der Universität Hamburg kindly loaned materials and facilitated two visits to the museum to examine materials.

Various people collected material for me or made gifts of material, sent specimens for identification, identified material, allowed access to collections under their care, made loans of material, provided other useful data and general information, and exchanged ideas. These are listed below in alphabetical order with their affiliations at the time of their contribution (and apologies if any titles are missing):-

K. Abbasi, Gilan Fisheries Research Centre, Bandar Anzali, H. A. Abdolhay, Tehran, A. Abdoli, Fisheries Research Centre, Sari and Gorgan University of Agricultural Sciences and Natural Resources, S. Abdolmalaki, Gilan Fisheries Research Centre, Bandar Anzali, Dr. T. Abe, University Museum, University of Tokyo, Dr. M. Abedi, Savadkooh University, H. Abyot, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Ahvaz, Dr. A. Adhami, Muze-ye Melli-ye Tarikh-e Tabi'i, Tehran, A. Afzali, Bandar Abbas, Dr. A. Akbary Pasand, University of Zabol, Zabol, A. Alamdari, Organization of the Environment, Shiraz, A. A. Al-Attar, Basrah University, A. W. Al-Hakim, University of Nottingham, L. A. J. Al-Hassan, School of Biological Sciences, University of Auckland, S. A. S. Al Hatimy, Oman Natural History Museum, Muscat, W. Al-Baharna, Directorate of Fisheries, Bahrein, Dr. N. M. Ali, Biological Research Centre, University of Baghdad, Dr. T. S. Ali, University of Basrah, S. Alinejad, Offshore Fisheries Research Centre, Chah Bahar, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, H. R. Alizadeh, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Tehran, A. R. Al-Jafery, Department of Hydrobiology, Baghdad, Dr. H. Alkahem, King Saud University, Riyadh, M. A. Al-Mukhtar, Fisheries Research Centre, Ahvaz, Prof. O. A. Amin, Arizona State University, Tempe, Dr. F. Andreone, Museo Regionale di Scienze Naturali, Torino, Dr. R. Arai, National Science Museum, Tokyo, G. Arbocco, Museo Civico di Storia Naturale "Giacomo Doria", Genova, Dr. J. D. Archibald, Yale University, Connecticut, Dr. N. B. Armantrout, Portland, Oregon, Dr. G. Arratia, University of Kansas, Lawrence, S. Asadollah, Isfahan University of Technology, A. Ashraf, Encyclopædia Iranica, Columbia University, New York, Dr. J. W. Atz, Department of Herpetology and Ichthyology, American Museum of Natural History, New York, Prof. S. Balik, Ege University, Izmir, Prof. E. Balletto, Istituto di Zoologia, Genova, G. A. C. Balma, Museo Civico di Storia Naturale, Carmagnola, Dr. K. Banister, Fish Section, British Museum (Natural History), London, A. J. Bardhun, Shiraz, D. M. Bartley, Food and Agriculture Organization, Rome, Dr. V. V. Barsukov, Zoological Institute, Academy of Sciences, Leningrad, M. L. Bauchot, Laboratoire d'Ichtyologie générale et appliquée, Muséum National d'Histoire Naturelle, Paris, R. Beck, COFAD GmbH, Tutzing, Dr. W. C. Beckman, Opelousas, Louisiana, Dr. A. Ben-Tuvia, Hebrew University of Jerusalem, Dr. M. Berberian, Uinversity of Cambridge, Dr. P. Berrebi, Université Montpellier, Dr. A. D. Berrie, Freshwater Biological Association, Wareham, Dr. E. Bertelsen, Zoologisk Museum, Copenhagen, Prof. Dr. P. G. Bianco, Universita degli Studi di l'Aquila, K. L. Bist, Government Postgraduate College, Gopeshwar, J. Bohlen, Academy of Sciences, Libechov, Dr. J. E. Böhlke, Academy of Natural Sciences, Philadelphia, Dr. A. H. Bornbusch, Duke University, Durham, Dr. J. Briggs, King Faisal university, Dammam, Dr. K. E. Carpenter, Food and Agriculture Organization, Rome, L. A. Cloutier, Department of the Environment, Tehran, Dr. D. Coffey, Pahlavi University, Shiraz, Dr. M. J. Collares-Pereira, Museu Bocage, Lisbon, Dr. J. T. Collins, Museum of Natural History, University of Kansas, Lawrence, Dr. L. J. V. Compagno, J. L. B. Smith Institute of Ichthyology, Grahamstown, Dr. B. B. Collette, National Museum of Natural History, Washington, G. H. Copp, Centre for Environment, Fisheries and Aquaculture Science, Lowestoft, Dr. L. Cornwallis, Oxford, A. S. Creighton, Division of Fishes, Museum of Zoology, University of Michigan, Ann Arbor, Dr. E. J. Crossman, Department of Ichthyology and Herpetology, Royal Ontario Museum, Toronto, E. L. Daniel, Encyclopædia Iranica, Columbia University, New York, F. Darvishi, Mazandaran, S. Deeb, American University of Lebanon, Beirut, S. Dehqan-Mediseh, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Ahvaz, Dr. G. B. Delmastro, Museo Civico di Storia Naturale, Carmagnola, M. Desoutter, Laboratoire d'Ichtyologie générale et appliquée, Museum National d'Histoire Naturelle, Paris, Dr. M. M. Dick, Museum of Comparative Zoology, Harvard University, Cambridge, P. Dickinson, National Zoological Garden, Al Ain, Abu Dhabi, W. A. Dill, Davis, California, J. Dominique, Freshwater and River Ecology Reserach Unit, Villeurbane, Dr. P. Dugan, Penang, Malayasia, M. Doroudi, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Bandar-e Lengeh, Dr. J. D. Durand, ESA CNRS, Villeurbane, Dr. G. Ekingen, Veteriner Fakultesi, Elazig, O.Elter, Museo ed Istituto di Zoologia Sistematico, Universita di Torino, Dr. B. Elvira, Ministerio de Agricultura y Pesca, Madrid, G. El Zein, Université Libanaise, Ksara, Dr. F. Erk'akan, Hacettepe University, Ankara, Dr. W. N. Eschmeyer, Department of Ichthyology, California Academy of Sciences, San Francisco, Gh. Eskandary, Fisheries Research Centre, Jahad-e Sazandegi, Ahvaz, Dr. H. R. Esmaeili, Shiraz University, D. Evans, IUCN, Cambridge, K. Evans, Pahlavi University, Shiraz, K. Fakhro, Directorate of Fisheries, Bahrein, R. Fatemi, Tehran, Dr. A. M. Fazel, Natural Resources Faculty, Tehran University, Karaj and Natural History Museum, Department of the Environment, Tehran, , H. Fazly, Fereydun Kenar, Mazandaran, R. F. Field, Muscat, Dr. E. Firouz, Tehran, Dr. W. Fischer, Food and Agriculture Organization, Rome, J. Fitzpatrick, Food and Agriculture Organization, Rome, Dr. R. Fricke, Staatliches Museum für Naturkunde in Stuttgart, P. A. M. Gaemers, Rijksmuseum van Geologie en Mineralogie, Leiden, M. D. Gallagher, Oman Natural History Museum, Muscat, M. Geerts, Swalmen, The Netherlands, Prof. Dr. R. Geldiay, Ege University, Izmir, Dr. C. George, Union College, Schenectady, Dr. H. Ghadirnejad, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Tehran, A. Ghamoosi, Shahid Beheshti University, Tehran, S. M. Ghasempouri, Tarbiat Modares University, Noor, Dr. D. I. Gibson, British Museum (Natural History), London, D. Golani, Zoological Museum, Hebrew University of Jerusalem, Dr. M. Goren, Tel Aviv University, S. Gorgin, Shiraz, Dr. B. Groombridge, UNEP World Conservation Monitoring Centre, Cambridge, Dr. S. H. Gruber, University of Miami, J. M. Gunn, University of Ottawa, R. Haas, California State University, Fresno, M. Hafezieh, Research Centre for Natural Resources and Animal Husbandry, Jahad-e Sazandegi, Shiraz, Dr. J. Halpern, Pahlavi University, Shiraz, Dr. K. E. Hartel, Museum of Comparative Zoology, Harvard University, Cambridge, S. S. Hasan, University of Basrah, Dr. M. R. Hassannia, Jahad-e Sazandegi, Tehran, M. R. Hemami, Isfahan University of Technology, D. M. Herdson, The Laboratory, Plymouth, E. Holm, Department of Ichthyology and Herpetology, Royal Ontario Museum, Toronto, Dr. R. A. Hinrichsen, Shad Foundation, Seattle, A.-M. Hodges, Fish Section, British Museum (Natural History), London, M. L. Holloway, Fish Section, British Museum (Natural History), London, L. Honarmond, University of Tehran, Dr. J. Holčík, Institute of Zoology, Slovak Academy of Sciences, Bratislava, Drs. F. and Sh. Hosseinie, Shiraz University, Dr. C. Hubbs, University of Texas, Austin, Dr. J. Huber, Muséum National d'Histoire Naturelle, Paris, J. Hull, University Museum, Oxford University, Dr. N. A. Hussain, Marine Science Centre, University of Basrah, Ch. Izadi, Research Centre for Natural Resources and Animal Husbandry, Jahad-e Sazandegi, Shiraz, Gh. Izadpanahi, Dr. B. Jalali, ABZIGOSTAR, Tehran, Dr. S. Jahromi, Pahlavi University, Shiraz, Dr. S. Jamili, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Tehran, Gh. A. Jasimi, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Ahvaz, Dr. M. N. Javed, Government College, Lahore, Dr. K. C. Jayaram, Zoological Survey of India, Calcutta, K. Jazebizadeh, Iranian Fisheries Research and Training Organization, Ahvaz, Dr. J. B. Jensen, Pahlavi University, Shiraz, Dr. R. K. Johnson, Field Museum of Natural History, Chicago, W. J. Jones, Al Ain, U.A.E., Dr. H. G. Kami, University of Tehran, J. M. Kapetsky, Food and Agriculture Organization, Rome, Dr. M. H. Karim Koshteh, University of Guelph, Dr. M. Kasparek, Kasparek Verlag, Heidelberg, Dr. E. J. Keall, Royal Ontario Museum, Toronto, Dr. A. Keyvanfar, Centre national de Transfusion sanguine-Institut, Paris, Dr. G. Khalaf, Lebanese University, Mansourieh-el-Metn, Dr. N. R. Khamees, University of Basrah, S. Khera, Punjab University, Chandigarh, A. Khodady, Shahid Chamran University, Ahvaz, Prof. Dr. R. Kinzelbach, Zoologisches Institut, Darmstadt, Dr. W. Klausewitz, Forschungsintitut Senckenberg, Frankfurt, Dr. W. L. Klawe, Inter-American Tropical Tuna Commission, Scripps Institution of Oceanography, La Jolla, Dr. M. Kottelat, Zoologsiches Staatsammlung, Munich, Dr. A. Kownacki, Laboratory of Water Biology, Polish Academy of Sciences, Krakow, Dr. S. O. Kullander, Swedish Museum of Natural History, Stockholm, Dr. K. Kuronuma, Tokyo University of Fisheries, Dr. M. Kuru, Hacettepe University, Ankara, P. Lamothe, Hydro Québec, Montréal, Dr. K. J. Lazara, US Merchant Marine Academy, Kings Point, New York, A. Lealmonfared, Shahid Beheshti University, Tehran, Dr. R. E. Lee, Pahlavi University, Shiraz, Dr. K. E. Limburg, State University of New York, Syracuse, Dr. R. Littman, University of Hawaii, Honolulu, Prof. Dr. H. Loffler, Vienna, R. Lolea, Gorgan University, J. Long, Department of Fisheries and Wildlife, Oregon State University, Corvallis, O. Lucanus, Montreal, Dr.Mabee, Department of Zoology, Duke University, Durham, A. A. Mahdi, University of Basrah, A. Mahjoor Azad, Shahid Beheshti University, Tehran, Dr. P. S. Maitland, Institute of Terrestrial Ecology, Edinburgh, Dr. H. Malicky, Biologische Station Lunz, L. Maltz, Tel Aviv University, J. Mansoori, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Ahvaz, J. Gh. Marammazi, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Ahvaz, R. Martino, American Killifish Association, Dr. M. Masoumian, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Tehran, Dr. A. Matinfar, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Tehran, Y. Mayahi, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Ahvaz, Dr. R. L. Mayden, Department of Biological Sciences, University of Alabama, Tuscaloosa, J. J. McAniff, National Underwater Accident Center, University of Rhode Island, Kingston, M. McDavitt, Alexandria, Virginia, S. Mickleburgh, Fauna and Flora Preservation Society, London, H.Meeus, Belgische Killifish Vereniging, Wommelgen, R. Mehrani, Lorestan Research Centre of Natural Resources and Animal Science, Khorramabad, Dr. A. G. K. Menon, Zoological Survey of India, Calcutta, Dr. S. N. Messieh, UNDP, Abu Dhabi, Dr. F. T. Mhaisen, University of Baghdad, Dr. A. Miller, Royal Botanic Garden, Edinburgh, I. D. Miller, United States-Saudi Arabian Joint Commission, New York, Dr. P. Miller, University of Bristol, Dr. R. R. Miller, Division of Fishes, Museum of Zoology, University of Michigan, Ann Arbor, Dr. A. A. Mirhosseyni, National Natural History Museum, Baghdad, Dr. M. R. Mirza, Lahore, A. Mobaraki, Department of the Environment, Tehran, M. R. Mohaghegh, Tehran, M. Mohammadi, Gorgan Agricultural and Natural Resources University, Dr. S. Moini, Department of the Environment, Tehran, Dr. B. Mokhayer, University of Tehran, Dr. K. Molnár, Veterinary Medical Research Institute, Hungarian Academy of Sciences, Budapest, Dr. F. Moravec, Institute of Parasitology, Czechoslovak Academy of Sciences, Prague, E. Morin, SOGREAH, Echirolles, Dr. E. O. Murdy, Bureau of Oceans and International Environmental and Scientific Affairs, Washington, Dr. G. S. Myers, Scotts Valley, California, M. Naderi, Mazandaran Fishery Research Centre, Sari, S. Naem, Faculty of Veterinary Medicine, Urmia University, A. Nasrollahzadeh, Gilan, Prof. Dr. C. M. Naumann, Universität Bielefeld, Dr. S. Nazeeri, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Khorramabad, R. B. Nehring, Department of the Environment, Tehran, N. Niameymandi, Persian Gulf Fisheries Research Centre, Bushehr, Dr. H. Nijssen, Instituut voor Taxonomisch Zoölogie, Zoölogisch Museum, Universiteit van Amsterdam, M. Nikpaey, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Ahvaz, N. Nouri, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Tehran, Dr. O. Oliva, Charles University, Prague, Dr. H.-J. Paepke, Museum für Naturkunde der Humboldt-Universität, Berlin, Dr. A. Paltrinieri, World Health Organization, Muscat, F. Papahn, Shahid Chamran University, Ahvaz, Dr. L. R. Parenti, National Museum of Natural History, Washington, J. Parkinson, Edmonton, A. Parsamanesh, Iranian Fisheries Research and Training Organization, Ahvaz, D. Peck, IUCN, Gland, T. Petr, Food and Agriculture Organization, Rome, H. Piri Zirkohy, Gilan Fisheries Research Centre, Bandar Anzali, Dr. E. P. Pister, Desert Fishes Council, Bishop, California, S. P. Platania, Colorado State University, Fort Collins, T. Plosch, Ganderkesee, L. Podshadley, Department of Ichthyology, California Academy of Sciences, San Francisco, Dr. M. Pourgholam, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Sari, M. Price, Division of Fishes, Museum of Zoology, University of Michigan, Ann Arbor, Dr. G. S. Proudlove, Department of Environmental Biology, University of Manchester, T. A. Qureshi, Technical Institute for Agriculture, Amara, M. Rabbaniha, Persian Gulf Fisheries Research Centre, Bushehr, Dr. H. Rahimian, University of Tehran, Dr. M. Ramin, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Tehran, F. M. Razi, Nature and Wildlife Museum, Tehran, Dr. W. J. Rainboth, University of California, Los Angeles, R. W. Redding, Museum of Zoology, University of Michigan, Ann Arbor, D. Rees, BBC, London, Dr. K. Relyea, Kuwait Institute for Scientific Research, H. Rezai, Tehran, Dr. S. Rezvani Gilkolaei, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Tehran, S. Richards, Murray, Utah, Dr. T. R. Roberts, Kasetsart University, Bangkok, Dr. I. Rostami, Shahid Chamran University, Ahvaz, B. Saadallah, Iraq Natural History Museum, Baghdad, M. A. G. Saadati, Department of the Environment, Mashhad, H. Saadoni, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Ahvaz, H. R. A. Sabet, Iranian Fisheries Research and Training Organization, Tehran, A. R. Saeed, University of Kerman, E. Saderigh-Nejad Massouleh, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Khorramabad, H. Safikhani, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Ahvaz, Dr. A. Salnikov, Institute of Zoology, Academy of Sciences, Ashkhabad, Dr. A. Samaie, Muse-ye Melli-ye Tarikh-e Tabi'i, Tehran, B. Sanford, Montrose, Colorado and Port Ludlow, Washington, Dr. A. Sanyal, Zoological Survey of India, Calcutta, Dr. M. Sarieyyüpoglu, Firat Üniversitesi, Elazig, Dr. A. Savari, Faculty of Oceanography, Shahid Chamran University, Ahvaz, M. Sayfali, Shahid Beheshti University, Tehran, D. A. Scott, Dursley, Gloucestershire, Dr. D. E. Sergeant, Arctic Biological Station, Ste-Anne de Bellevue, Quebec, Gh. Shakhiba, Iranian Fisheries Research and Training Organization, Ahvaz, A. J. Shams, Directorate of Fisheries, Bahrein, Dr. I. Sharifpour, Iranian Fisheries Research and Training Organization, Ahvaz, J. W. Sherman, Academy of Natural Sciences, Philadelphia, Dr. A. Shiralipour, Pahlavi University, Shiraz, Dr. I. Q. Siddiqui, King Faisal University, Al Hasa, Dr. P. Skelton, Fish Section, British Museum (Natural History), London, Dr. G. R. Smith, Museum of Zoology, University of Michigan, Ann Arbor, Dr. W. F. Smith-Vaniz, Academy of Sciences, Philadelphia, M. Soleymani, Green Front of Iran, Tehran, N. Statman, Dr. A. N. Svetovidov, Zoological Institute, Academy of Sciences, Leningrad, Dr. C. C. Swift, Natural History Museum of Los Angeles County, Dr. F. Terofal, Zoologische Sammlung des Bayreischen Staates, Munich, M. V. Tofighi, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Tehran, A. Torfi, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Ahvaz, Dr. W. Torke, Institut fur Urgeschichte, Tübingen, Dr. E. Tortonese, Museo Civico di Storia Naturale, Genova, Dr. R. A. Travers, Fish Section, British Museum (Natural History), London, R. G. Tuck, Muze-ye Melli-ye Tarikh-e Tabi'i, Tehran, Dr. H. Türkmen, Istanbul Üniversitesi, Dr. E. Unlu, University of Dicle, Diyarbakir, Dr. I. Unsal, Istanbul Üniversitesi, T. Valinasab, Fisheries Research and Training Organization, Jahad-e Sazandegi, Tehran, Dr. J. Valiallahi, Tarbiat-e Modarres, Noor, >W. van Neer, Royal Museum of Central Africa, Tervuren, Prof. Dr. R. Victor, Sultan Qaboos University, Muscat, Prof. Dr. W. Villwock, Zoologisches Institut und Zoologisches Museum, Hamburg, Dr. V. D. Vladykov, University of Ottawa, A. Vosughi, Iranian Fisheries Research and Training Organization, Jahad-e Sazandegi, Tehran, B. Waaland, Pahlavi University, Shiraz, P. Walczak, Department of the Environment, Tehran, Dr. B. A. Whitton, University of Durham, Dr. R. Winterbottom, Department of Ichthyology and Herpetology, Royal Ontario Museum, Toronto, Dr. G. H. Wossughi, University of Tehran, Dr. T. C. Young, Royal Ontario Museum, Toronto, M. Zapater, Zaragoza, A. R. Zeanaie, Payam-e Noor University, Bandar Abbas.

Individual Iranians, too numerous to mention here, kindly enunciated carefully and repeatedly Farsi fish names for my cloth ear. However it would be remiss not to mention staff at the Iranian Fisheries Research and Training Organization, Ahvaz including N. Najafpour, Gh. Marammazi, Gh. Eskandari, and M. A. Al-Mukhtar, as well as E. Firouz, Tehran, B. Kiabi and A. Abdoli, Gorgan Agricultural and Natural Resources University, and Y. Keivany, University of Alberta, Edmonton.

And finally I must thank my wife Sylvie and son Nicholas for supporting me in my obsession with fishes from Iran and Nick for constructing the index page for this website and linking it to the internet.

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© Brian W. Coad (www.briancoad.com)