The nominal species originally described from Iran are summarised below by family, by year of description and then alphabetically. Some taxa have no type material (e.g. Eugen Keyserling did not bring specimens back to Europe) or type material has not been located despite a thorough search and is marked with an asterisk (*). Types whose existence is uncertain are indicated by a question mark (?). Type material examined by me is indicated by a plus sign (+) and type material not seen by me by an octothorpe (#). The current allocation of taxa is given in parentheses, those without comment being valid; material of uncertain status is marked with a tilde (~).

Details on these fishes can be found in the Species Accounts, where other type material examined for its importance in studying the Iranian ichthyofauna is listed. See also Eschmeyer's Catalog of Fishes (www.calacademy.org/research/ichthyology/catalog/).

Family Clupeidae

# Caspialosa caspia var. persica Il'in, 1927 (= Alosa caspia (Eichwald, 1838))

# Caspialosa knipowitschi Il'in, 1927 (= Alosa caspia (Eichwald, 1838))

# Clupea caspio-pontica var. grimmi Borodin, 1904 (= Alosa braschnikowii (Borodin, 1904))

Family Cyprinidae

*# Cyprinus persa Gmelin, 1774 (nomen nudum = Vimba vimba (Linnaeus, 1758))

*# Cyprinus bulatmai Hablizl, 1783 (= Barbus capito (Güldenstaedt, 1773))

?# Cyprinus chalybatus Pallas, 1814 (= Barbus capito (Güldenstaedt, 1773))

+ Alburnus capito Heckel, 1843 (= Alburnus mossulensis Heckel, 1843)

+ Chondrostoma aculeata Valenciennes in Cuvier and Valenciennes, 1844 (Capoeta aculeata Valenciennes in Cuvier and Valenciennes, 1844))

# Leuciscus albuloides Valenciennes in Cuvier and Valenciennes, 1844 (= Alburnus chalcoides (Güldenstaedt, 1772))

+ Leuciscus maxillaris Valenciennes in Cuvier and Valenciennes, 1844 (= Alburnus mossulensis Heckel, 1843)

+ Alburnus caudimacula Heckel, 1849 (= Alburnus mossulensis Heckel, 1843)

+ Alburnus iblis Heckel, 1849 (= Alburnus mossulensis Heckel, 1843)

+ Alburnus megacephalus Heckel, 1849 (= Alburnus mossulensis Heckel, 1843)

+ Alburnus schejtan Heckel, 1849 (= Alburnus mossulensis Heckel, 1843)

+ Barbus barbulus Heckel, 1849

+ Cyprinion tenuiradius Heckel, 1849

+ Discognathus crenulatus Heckel, 1849 (= Garra rufa (Heckel, 1843))

+ Scaphiodon amir Heckel, 1849 (Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842))

+ Scaphiodon macrolepis Heckel, 1849 (= Capoeta aculeata Valenciennes in Cuvier and Valenciennes, 1844))

+ Scaphiodon niger Heckel, 1849 (Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842))

+ Scaphiodon saadii Heckel, 1849 (Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842))

+ Systomus albus var. alpina Heckel, 1849 (= Barbus luteus (Heckel, 1843))

*#~ Alburnus maculatus Keyserling, 1861

*# Scaphiodon chebisiensis Keyserling, 1861 (Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842))

*# Scaphiodon gracilis Keyserling, 1861 (= Capoeta capoeta (Güldenstaedt, 1773))

*# Scaphiodon rostratus Keyserling, 1861 (= Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842))

?# ~ Barbus miliaris De Filippi, 1863

+~ Alburnus doriae De Filippi, 1865

+ Schizothorax pelzami Kessler, 1870

+ Capoeta buhsei Kessler, 1877

+ Aspiostoma zarudnyi Nikol'skii, 1897 (= Schizothorax zarudnyi (Nikol'skii, 1897))

+ Capoeta fusca Nikol'skii, 1897

+ Capoeta gibbosa Nikol'skii, 1897 (= Capoeta capoeta (Güldenstaedt, 1773))

+ Capoeta nudiventris Nikol'skii, 1897 (= Capoeta fusca Nikol'skii, 1897)

+ Abramis urmianus Günther, 1899 (= Acanthalburnus urmianus (Günther, 1899))

+ Barbus bampurensis Nikol'skii, 1899 (= Cyprinion watsoni (Day, 1872))

+ Cyprinion kirmanense Nikol'skii, 1899 (= Cyprinion watsoni (Day, 1872))

+ Gobio persa Günther, 1899 (= Romanogobio persus (Günther, 1899))

+ Leuciscus gaderanus Günther, 1899 (= Petroleuciscus ulanus (Günther, 1899))

+ Leuciscus ulanus Günther, 1899 (= Petroleuciscus ulanus (Günther, 1899))

+ Cirrhina afghana var. nikolskii Berg, 1905 (= Cyprinion watsoni (Day, 1872))

+ Discognathus rossicus var. nudiventris Berg, 1905 (= Garra rossica (Nikol'skii, 1900))

+ Scaphiodon macmahoni Regan, 1906 (= Cyprinion watsoni (Day, 1872))

+ Garra persica Berg, 1913

+ Discognathus adiscus Annandale, 1919 (= Crossocheilus latius (Hamilton-Buchanan, 1822))

+ Discognathus phryne Annandale, 1919 (= Garra rossica (Nikol'skii, 1900))

+ Alburnus atropatenae Berg, 1925 (= Alburnus atropatenae Berg, 1925)

?# Alburnus alburnus hohenackeri natio persicus Petrov, 1926 (= Alburnus alburnus (Linnaeus, 1758))

?# Alburnus pseudospirlinus Petrov, 1926 (= Alburnus alburnus (Linnaeus, 1758))

?# Alburnus striatus Petrov, 1926 (= Alburnus alburnus (Linnaeus, 1758))

?# Rutilus rutilus caspicus natio knipowitschi Pravdin, 1927

+ Barbus baschakirdi Holly, 1929 (= Cyprinion watsoni (Day, 1872))

*# Barbus kessleri Derzhavin, 1929 (Barbus miliaris De Filippi, 1863)

*# Varicorhinus bergi Derzhavin, 1929 (= Capoeta aculeata Valenciennes in Cuvier and Valenciennes, 1844)

*# Varicorhinus nikolskii Derzhavin, 1929 (= Capoeta buhsei Kessler, 1877)

+ Barilius mesopotamicus Berg, 1932

?# Leuciscus cephalus orientalis natio ardebilicus Barach, 1934

+ Iranocypris typhlops Bruun and Kaiser, 1944

+ Chalcalburnus chalcoides iranicus Svetovidov, 1945 (= Alburnus chalcoides (Güldenstaedt, 1772))

+ Garra rufa gymnothorax Berg, 1949 (Garra rufa (Heckel, 1843))

+ Oreinus anjac Fowler and Steinitz, 1956 (Schizothorax zarudnyi (Nikol'skii, 1897))

+ Schizothorax schumacheri Fowler and Steinitz, 1956 (= Schizothorax intermedius McClelland, 1842)

+ Capoeta barroisi persica Karaman, 1969

+ Schizothorax pelzami iranicus Karaman, 1969 (= Schizothorax pelzami Kessler, 1870)

+ Pseudophoxinus persidis Coad, 1981 (Petroleuciscus persidis (Coad, 1982))

?# Barbus parieschanica Wossughi, Khoshzahmat and Etemadfar, 1982 (= Barbus luteus (Heckel, 1843))

+ Capoeta barroisi mandica Bianco and Banarescu, 1982

+ Capoeta capoeta intermedia Bianco and Banarescu, 1982

+ Chondrostoma cyri orientalis Bianco and Banarescu, 1982 (= Chondrostoma regium (Heckel, 1843))

+ Schizocypris altidorsalis Bianco and Banarescu, 1982

+ Barbus sublimus Coad and Najafpour, 1997

Family Cobitidae

+ Acanthopsis linea Heckel, 1849 (= Cobitis linea (Heckel, 1849))

# Cobitis aurata De Filippi, 1863 (= Sabanejewia aurata (De Filippi, 1863))

Family Balitoridae

+ Cobitis persa Heckel, 1849 (= Barbatula persa (Heckel, 1849))

+ Nemacheilus bampurensis Nikol'skii, 1899 (= Schistura bampurensis (Nikol'skii, 1899)

+ Nemacheilus sargadensis Nikol'skii, 1899 (= Schistura sargadensis (Nikol'skii, 1899)

+ Nemachilus rhadinaeus Regan, 1906 (= Paracobitis rhadinaea (Regan, 1906))

+ Nemachilus macmahoni Chaudhuri, 1909 (= Paracobitis rhadinaea (Regan, 1906))

+ Nemachilus bergianus Derzhavin, 1934 (= Barbatula bergiana (Derzhavin, 1934))

+ Noemacheilus kermanshahensis Bănărescu and Nalbant, 1967 (= Barbatula kermanshahensis (Bănărescu and Nalbant, 1967))

+ Nemacheilus smithi Greenwood, 1976 (= Paracobitis smithi Greenwood, 1976))

+ Orthrias farsicus Nalbant and Bianco, 1998 (= Barbatula farsica (Nalbant and Bianco, 1998))

+ Paracobitis iranica Nalbant and Bianco, 1998

+ Paracobitis vignai Nalbant and Bianco, 1998

+ Schistura nielseni Nalbant and Bianco, 1998

? Seminemacheilus tongiorgii Nalbant and Bianco, 1998

Family Sisoridae

+ Glyptosternum kurdistanicum Berg, 1931 (= Glyptothorax kurdistanicus (Berg, 1931))

+ Glyptothorax silviae Coad, 1981

Family Cyprinodontidae

+ Lebias crystallodon Heckel, 1849 (= Aphanius sophiae (Heckel, 1849))

+ Lebias punctatus Heckel, 1849 (= Aphanius sophiae (Heckel, 1849))

+ Lebias sophiae Heckel, 1849 (= Aphanius sophiae (Heckel, 1849))

#~ Cyprinodon blanfordii Jenkins, 1910

# Cyprinodon persicus Jenkins, 1910 (= Aphanius persicus (Jenkins, 1910))

#~ Cyprinodon pluristriatus Jenkins, 1910

+ Cyprinodon ginaonis Holly, 1929 (= Aphanius ginaonis (Holly, 1929)

+ Aphanius vladykovi Coad, 1988

+ Aphanius isfahanensis Hrbek, Keivany and Coad, 2007

Family Cichlidae

+ Iranocichla hormuzensis Coad, 1982

Family Mugilidae

+ Liza abu zarudnyi Berg, 1949

Family Gobiidae